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  <front>
    <journal-meta>
      <journal-id journal-id-type="publisher-id">102</journal-id>
      <journal-id journal-id-type="index">urn:lsid:arphahub.com:pub:73abe0ce-d97c-5d7c-bee5-b8e6e6fe6a17</journal-id>
      <journal-title-group>
        <journal-title xml:lang="en">ARPHA Preprints</journal-title>
        <abbrev-journal-title xml:lang="en">preprints</abbrev-journal-title>
      </journal-title-group>
      <publisher>
        <publisher-name>Pensoft Publishers</publisher-name>
      </publisher>
    </journal-meta>
    <article-meta>
      <article-id pub-id-type="doi">10.3897/arphapreprints.e120539</article-id>
      <article-id pub-id-type="publisher-id">120539</article-id>
      <article-categories>
        <subj-group subj-group-type="heading">
          <subject>Research Article</subject>
        </subj-group>
        <subj-group subj-group-type="biological_taxon">
          <subject>Amphibia</subject>
        </subj-group>
        <subj-group subj-group-type="scientific_subject">
          <subject>Biodiversity &amp; Conservation</subject>
          <subject>Biogeography</subject>
        </subj-group>
        <subj-group subj-group-type="geographical_area">
          <subject>Asia</subject>
        </subj-group>
      </article-categories>
      <title-group>
        <article-title>Range extension and expanded description of <italic>Micryletta hekouensis</italic></article-title>
      </title-group>
      <contrib-group content-type="authors">
        <contrib contrib-type="author" corresp="yes">
          <name name-style="western">
            <surname>Tan</surname>
            <given-names>Nguyen Van</given-names>
          </name>
          <email xlink:type="simple">tan.sifasv@gmail.com</email>
          <uri content-type="orcid">https://orcid.org/0000-0001-5413-968X</uri>
          <xref ref-type="aff" rid="A1">1</xref>
        </contrib>
        <contrib contrib-type="author" corresp="no">
          <name name-style="western">
            <surname>Liu</surname>
            <given-names>Shuo</given-names>
          </name>
          <uri content-type="orcid">https://orcid.org/0000-0001-7825-3006</uri>
          <xref ref-type="aff" rid="A2">2</xref>
        </contrib>
        <contrib contrib-type="author" corresp="no">
          <name name-style="western">
            <surname>Tran</surname>
            <given-names>Vy</given-names>
          </name>
          <xref ref-type="aff" rid="A3">3</xref>
        </contrib>
        <contrib contrib-type="author" corresp="no">
          <name name-style="western">
            <surname>Tran</surname>
            <given-names>Thinh G.</given-names>
          </name>
          <uri content-type="orcid">https://orcid.org/0009-0009-7386-6503</uri>
          <xref ref-type="aff" rid="A4">4</xref>
        </contrib>
        <contrib contrib-type="author" corresp="no">
          <name name-style="western">
            <surname>Trofimets</surname>
            <given-names>Alexei V.</given-names>
          </name>
          <xref ref-type="aff" rid="A5">5</xref>
        </contrib>
        <contrib contrib-type="author" corresp="yes">
          <name name-style="western">
            <surname>Dau</surname>
            <given-names>Vinh</given-names>
          </name>
          <email xlink:type="simple">dauquangvinhna@hdu.edu.vn</email>
          <xref ref-type="aff" rid="A6">6</xref>
        </contrib>
        <contrib contrib-type="author" corresp="yes">
          <name name-style="western">
            <surname>Poyarkov</surname>
            <given-names>Nikolay</given-names>
          </name>
          <email xlink:type="simple">n.poyarkov@gmail.com</email>
          <uri content-type="orcid">https://orcid.org/0000-0002-7576-2283</uri>
          <xref ref-type="aff" rid="A7">7</xref>
        </contrib>
      </contrib-group>
      <aff id="A1">
        <label>1</label>
        <addr-line content-type="verbatim">Snake identification and First aid for snakebites in Vietnam, Ninh Binh, Vietnam</addr-line>
        <institution>Snake identification and First aid for snakebites in Vietnam</institution>
        <addr-line content-type="city">Ninh Binh</addr-line>
        <country>Vietnam</country>
      </aff>
      <aff id="A2">
        <label>2</label>
        <addr-line content-type="verbatim">Kunming Natural History Museum of Zoology, Kunming Institute of Zoology, Chinese Academy of Sciences, 32 Jiaochang Donglu, Kunming, Yunnan 650223, China, Kunming, China</addr-line>
        <institution>Kunming Natural History Museum of Zoology, Kunming Institute of Zoology, Chinese Academy of Sciences, 32 Jiaochang Donglu, Kunming, Yunnan 650223, China</institution>
        <addr-line content-type="city">Kunming</addr-line>
        <country>China</country>
      </aff>
      <aff id="A3">
        <label>3</label>
        <addr-line content-type="verbatim">Institute of Tropical Biology, Vietnam Academy of Science and Technology, 85 Tran Quoc Toan St., District 3, Ho Chi Minh City, Vietnam, Ho Chi Minh, Vietnam</addr-line>
        <institution>Institute of Tropical Biology, Vietnam Academy of Science and Technology, 85 Tran Quoc Toan St., District 3, Ho Chi Minh City, Vietnam</institution>
        <addr-line content-type="city">Ho Chi Minh</addr-line>
        <country>Vietnam</country>
      </aff>
      <aff id="A4">
        <label>4</label>
        <addr-line content-type="verbatim">Department of Ecology and Evolutionary Biology, Faculty of Biology and Biotechnology, University of Science, Ho Chi Minh City, 227 Nguyen Van Cu Street, District 5, Ho Chi Minh City, Vietnam, Ho Chi Minh, Vietnam</addr-line>
        <institution>Department of Ecology and Evolutionary Biology, Faculty of Biology and Biotechnology, University of Science, Ho Chi Minh City, 227 Nguyen Van Cu Street, District 5, Ho Chi Minh City, Vietnam</institution>
        <addr-line content-type="city">Ho Chi Minh</addr-line>
        <country>Vietnam</country>
      </aff>
      <aff id="A5">
        <label>5</label>
        <addr-line content-type="verbatim">Department of Vertebrate Zoology, Lomonosov Moscow State University, Leninskiye Gory, GSP–1, Moscow 119991, Russia, Moscow, Russia</addr-line>
        <institution>Department of Vertebrate Zoology, Lomonosov Moscow State University, Leninskiye Gory, GSP–1, Moscow 119991, Russia</institution>
        <addr-line content-type="city">Moscow</addr-line>
        <country>Russia</country>
      </aff>
      <aff id="A6">
        <label>6</label>
        <addr-line content-type="verbatim">Hong Duc University, 565 Quang Trung Street-Dong Ve Ward-Thanh Hoa City, Vietnam, Thanh Hoa, Vietnam</addr-line>
        <institution>Hong Duc University, 565 Quang Trung Street-Dong Ve Ward-Thanh Hoa City, Vietnam</institution>
        <addr-line content-type="city">Thanh Hoa</addr-line>
        <country>Vietnam</country>
      </aff>
      <aff id="A7">
        <label>7</label>
        <addr-line content-type="verbatim">Joint Vietnam - Russia Tropical Science and Technology Research Centre, 63 Nguyen Van Huyen Road, Nghia Do, Cau Giay, Hanoi, Vietnam, Ha Noi, Vietnam</addr-line>
        <institution>Joint Vietnam - Russia Tropical Science and Technology Research Centre, 63 Nguyen Van Huyen Road, Nghia Do, Cau Giay, Hanoi, Vietnam</institution>
        <addr-line content-type="city">Ha Noi</addr-line>
        <country>Vietnam</country>
      </aff>
      <author-notes>
        <fn fn-type="corresp">
          <p>Corresponding authors: Nguyen Van Tan (<email xlink:type="simple">tan.sifasv@gmail.com</email>), Vinh Dau (<email xlink:type="simple">dauquangvinhna@hdu.edu.vn</email>), Nikolay Poyarkov (<email xlink:type="simple">n.poyarkov@gmail.com</email>).</p>
        </fn>
        <fn fn-type="edited-by">
          <p>Academic editor: </p>
        </fn>
      </author-notes>
      <pub-date pub-type="collection">
        <year>2024</year>
      </pub-date>
      <pub-date pub-type="epub">
        <day>09</day>
        <month>02</month>
        <year>2024</year>
      </pub-date>
      <volume>5</volume>
      <uri content-type="arpha" xlink:href="http://openbiodiv.net/9A4B3E66-825D-5A01-96AD-EDF2193F2E90">9A4B3E66-825D-5A01-96AD-EDF2193F2E90</uri>
      <history>
        <date date-type="received">
          <day>08</day>
          <month>02</month>
          <year>2024</year>
        </date>
        <date date-type="accepted">
          <day>08</day>
          <month>02</month>
          <year>2024</year>
        </date>
      </history>
      <permissions>
        <copyright-statement>Nguyen Van Tan, Shuo Liu, Vy Tran, Thinh G. Tran, Alexei V. Trofimets, Vinh Dau, Nikolay Poyarkov</copyright-statement>
        <license license-type="creative-commons-attribution" xlink:href="http://creativecommons.org/licenses/by/4.0/" xlink:type="simple">
          <license-p>This is an open access preprint distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.</license-p>
        </license>
      </permissions>
      <abstract>
        <label>Abstract</label>
        <p>The Paddy frog species <italic>Micryletta hekouensis</italic><bold> </bold>was described based on only two specimens from Nanxi Village, Nanxi Town, Hekou County, Honghe Prefecture, Yunnan Province, China. Herein, we report on new findings and a range extension of this species based on a re-examination of preserved specimens deposited in Duy Tan University (DTU) and Zoological Museum of Lomonosov Moscow State University (ZMMU) collected from Vietnam. All new specimens were previously identified as <italic>Micryletta</italic> cf. <italic>inornata </italic>or<italic> M</italic>.<italic> </italic>cf. <italic>steinegeri</italic>. Molecular analyses based on mitochondrial DNA supported the morphological findings. The newly identified specimens have a pairwise divergence of only 0.7% from those of the type series of <italic>M</italic>.<italic> </italic><italic>hekouensis </italic>(based on the 16s rRNA mtDNA gene). Based on the new information, we confirm the presence of <italic>M</italic>.<italic> </italic><italic>hekouensis</italic> in Vietnam and update the diagnostic characters of this species and distribution. We suggest the species should be considered as Near Threatened (NT) following the IUCN’s Red List categories. Further studies reassessing the populations of the <italic>Micryletta</italic> <italic>inornata </italic>complex in Indochinese Region (including: Vietnam, Laos, Cambodia, Thailand) are required.</p>
      </abstract>
    </article-meta>
  </front>
</article>
