MFS collected the data, served as authority on species identification, provide photograph of the specimens, analyzed the data, wrote, and revised the manuscript. BHBS collected the data, served as authority on species identification, provide photograph of the specimens and constructed the map figure. SAF, MIMAW, MAN, FHA, GK, NNS, NFN, MS-R, JA, TKL, NSX, MFA, SAR collected the data. ABA conceptualized the study design, analyzed the data and revised the manuscript.