Preprint
ARPHA Preprints
https://doi.org/10.3897/arphapreprints.e161758 (11 Jun 2025)
https://doi.org/10.3897/arphapreprints.e161758 (11 Jun 2025)
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ARPHA Preprints
doi:
10.3897/arphapreprints.e161758
First posted
11 Jun 2025
Authors
Ileana Herrera
- Corresponding author
Instituto Nacional de Biodiversidad (INABIO), Quito, Ecuador
Universidad Espiritu Santo (UEES), Samborondon, Ecuador
Centro de Investigaciones, Universidad Espíritu Santo, Samborondón, Ecuador, Samborondón, Ecuador
Centro de Investigaciones, Universidad Espíritu Santo, Samborondón, Ecuador, Samborondón, Ecuador

Universidad Nacional de Loja, Loja, Ecuador
Centro de Investigaciones, Universidad Espíritu Santo, Samborondón, Ecuador, Samborondón, Ecuador
Vrije Universiteit Brussel, Department of Biology, Brussels, Belgium
School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom
Centro de Investigaciones, Universidad Espíritu Santo, Samborondón, Ecuador, Samborondón, Ecuador
School of Biological, Earth and Environmental Sciences, University of New South Wales, Sydney, Australia
Instituto Nacional de Biodiversidad, Quito, Ecuador
Institut Botànic de Barcelona (IBB), CSIC-CMCNB, Barcelona, Spain
Universidad Estatal Amazónica, Puyo, Ecuador
Vanessa Lucia Lozano
- Corresponding author
Department of Agricultural Sciences, University of Sassari, Viale Italia 39, 07100 Sassari, Italy, Sassari, Italy
Departamento de Biología, Escuela Politécnica Nacional, Quito, Ecuador
Facultad de Ciencias de Medio Ambiente, Universidad Tecnológica Indoamérica, Quito, Ecuador
Centro de Investigaciones, Universidad Espíritu Santo, Samborondón, Ecuador, Samborondón, Ecuador
Universidad Nacional de Loja, Loja, Ecuador
Herbario Departamento de Biología, Roca Quito, Ecuador
School of Biological Sciences and Engineering, Yachay Tech University, Urcuquí, Ecuador
Missouri Botanical Garden, St. Louis, United States of America
Centro de Investigaciones, Universidad Espíritu Santo, Samborondón, Ecuador, Samborondón, Ecuador
Jordi López-Pujol
- Corresponding author
Botanic Institute of Barcelona (IBB, CSIC-ICUB), Barcelona, Spain
Conflict of interest
The authors have declared that no competing interests exist.
Supporting agencies
This study was partly supported by the Research Center of the Universidad Espíritu Santo (to IH), grant number 2024-ING-004, and by the Ministerio de Ciencia e Innovación, Spain (project PID2020-119163GB-I00 funded by MCIN/AEI/10.13039/501100011033) (to JL-P). VL was supported by the PON-AIM (Programma Operativo Nazionale ricerca e innovazione 2014–2020; ID AIM1897595-2 and ID DM1062AGR-IV6BRUNDU) and by the
University of Sassari (Italy) and the Autonomous Region of Sardinia - Legge regionale 7 agosto 2007, n.7 “Promozione della Ricerca Scientifica e dell’Innovazione Tecnologica in Sardegna”, “Mobilità Giovani Ricercatori (MGR)”.
This is an open access preprint distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.