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ARPHA Preprints
https://doi.org/10.3897/arphapreprints.e188569 (17 Feb 2026)
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ARPHA Preprints
doi: 10.3897/arphapreprints.e188569
First posted 17 Feb 2026
Authors
Yonglin Tan
Shandong Second Medical University, Weiping, China
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Jinxin Chu
Shandong Second Medical University, Weifang, China
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Hui Li
Weifang Institute of Technology, Weifang, China
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liangning Yin
Shandong Second Medical University, Weifang, China
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Xinrui Sun
Shandong Second Medical University, Weifang, China
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Dongyan Fu
Shandong Second Medical University, Weifang, China
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Weiping Lin - Corresponding author
Shandong Second Medical University, Weifang, China
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Hai Dian Hou - Corresponding author
Shandong Second Medical University, Weifang, China
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Author contributions

Jinxin Chu, Dianhai Hou and Weiping Lin conceived the original idea and designed the work; Yonglin Tan, Jinxin Chu and Hui Li performed the phylogenetic analyses; Yonglin Tan, Jinxin Chu, Hui Li, Liangning Yin, Xinrui Sun, and Dongyan Fu collected sequence data, and Yonglin Tan and Dianhai Hou wrote the paper and discussed with all authors.


Conflict of interest
The authors have declared that no competing interests exist.
Supporting agencies
Shandong Provincial Natural Science Foundation (No. ZR2020QC014), and the Doctoral Fund of Weifang Medical University (No. 02181801).
This is an open access preprint distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
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