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ARPHA Preprints
https://doi.org/10.3897/arphapreprints.e189735 (25 Feb 2026)
 
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ARPHA Preprints
doi: 10.3897/arphapreprints.e189735
First posted 25 Feb 2026
Authors
   
   
Rafael Marquina Blasco - Corresponding author
GIUV2016-303 Grup d'Investigació En Paleontologia de Vertebrats del Cenozoic PVC-GIUV, Departament de Botànica i Geologia, Àrea de Paleontologia, Universitat de València, Valencia, Spain, VALENCIA, Spain
Centro de Investigação em Ciências Geo-Espaciais (CICGE), Universidade do Porto, Vila Nova de Gaia, Portugal, Vila Nova de Gaia, Portugal
Museu Valencià d'Història Natural, Alginet, Spain
Institut Catalá de Paleoecologia Humana i Evolució Social (IPHES-CERCA), Tarragona, Spain
Articles by this author in:  CrossRef  |  PubMed  |  Google Scholar
Plinio Montoya
GIUV2016-303 Grup d'Investigació En Paleontologia de Vertebrats del Cenozoic PVC-GIUV, Departament de Botànica i Geologia, Àrea de Paleontologia, Universitat de València, Valencia, Spain, Valencia, Spain
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Francisco Javier Ruiz-Sánchez
GIUV2016-303 Grup d'Investigació En Paleontologia de Vertebrats del Cenozoic PVC-GIUV, Departament de Botànica i Geologia, Àrea de Paleontologia, Universitat de València, Valencia, Spain, VALENCIA, Spain
Museu Valencià d'Història Natural, Alginet, Spain
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Vicente D. Crespo
Departamento de Ciências da Terra, FCT-UNL Faculdade de Ciências E Tecnologia, G eoBioTec, Universidade Nova de Lisboa, Caparica, Portugal, Caparica, Portugal
Museu da Lourinhã, 9 Rua João Luis de Moura, 2530-158, Lourinhã, Portugal, Lourinhã, Portugal
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Author contributions
Rafael Marquina-Blasco: Conceptualisation, methodology, formal analysis, data curation, writing—original draft, writing—review and editing. Plinio Montoya: Conceptualisation, writing—review and editing, supervision, funding acquisition. Francisco Javier Ruiz-Sánchez: Conceptualisation, writing—review and editing, supervision, funding acquisition. Vicente D. Crespo: Conceptualisation, methodology, writing—original draft, writing—review and editing.  
Conflict of interest
The authors have declared that no competing interests exist.
Supporting agencies
CIUCSD -  Conselleria de Innovación, Universidades, Ciencia y Sociedad Digital, Generalitat Valenciana
SFRH/BPD/88207/2012 -  FCT POSTDOC
MCIU -  Ministerio de Ciencia, Innovación y Universidades
This is an open access preprint distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
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