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ARPHA Preprints
https://doi.org/10.3897/arphapreprints.e191104 (10 Mar 2026)
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ARPHA Preprints
doi: 10.3897/arphapreprints.e191104
First posted 10 Mar 2026
Authors
   
   
   
Christos Gkenas - Corresponding author
MARE - Centro de Ciências do Mar e do Ambiente, Lisboa, Portugal
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Nicholas Koutsikos
Hellenic Centre for Marine Research, Institute of Marine Biological Resources and Inland Waters, Athens, Greece
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Katelyn Lawson
Auburn University, Auburn, United States of America
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Filipe Ribeiro
MARE - Marine and Environmental Sciences Centre, Lisbon, Portugal
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Leonidas Vardakas
University of the Aegean, Mytilene, Greece
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Author contributions

CG and FR contributed to the study conception and design. Material preparation and data collection were performed by CG, NK, KL and LV. Data analysis and visualisation were performed by CG. CG and FR secured funding. The first draft of the manuscript was written by CG and all authors commented on previous versions of the manuscript. All authors read and approved the final manuscript.


Conflict of interest
The authors have declared that no competing interests exist.
This is an open access preprint distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
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