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ARPHA Preprints
https://doi.org/10.3897/arphapreprints.e194582 (07 Apr 2026)
Submitted to Frontiers of Biogeography
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ARPHA Preprints
doi: 10.3897/arphapreprints.e194582
First posted 07 Apr 2026
Authors
   
   
   
   
Ivan Nekhaev
Institute of Zoology, Almaty, Kazakhstan
Saint-Petersburg State University, Saint Petersburg, Russia
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Amina Omarova - Corresponding author
Institute of Zoology, Almaty, Kazakhstan
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Anel Ishayeva
Institute of Zoology, Almaty, Kazakhstan
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Author contributions

Ivan Nekhaev: conceptualization; field sampling; morphological analysis; phylogenetic analysis and species distribution modelling; preparation of the original draft. Amina Omarova: DNA extraction; polymerase chain reaction; sequence processing; molecular phylogenetic analysis. Anel Ishayeva: organisation of field expeditions; specimen collection.


Conflict of interest
The authors have declared that no competing interests exist.
Supporting agencies
Science Committee of the Ministry of Science and Higher Education of the Republic of Kazakhstan (Grant No. AP22783298)
This is an open access preprint distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
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