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  <front>
    <journal-meta>
      <journal-id journal-id-type="publisher-id">102</journal-id>
      <journal-id journal-id-type="index">urn:lsid:arphahub.com:pub:73abe0ce-d97c-5d7c-bee5-b8e6e6fe6a17</journal-id>
      <journal-title-group>
        <journal-title xml:lang="en">ARPHA Preprints</journal-title>
        <abbrev-journal-title xml:lang="en">preprints</abbrev-journal-title>
      </journal-title-group>
      <publisher>
        <publisher-name>Pensoft Publishers</publisher-name>
      </publisher>
    </journal-meta>
    <article-meta>
      <article-id pub-id-type="doi">10.3897/arphapreprints.e69207</article-id>
      <article-id pub-id-type="publisher-id">69207</article-id>
      <article-categories>
        <subj-group subj-group-type="heading">
          <subject>Research Article</subject>
        </subj-group>
        <subj-group subj-group-type="biological_taxon">
          <subject>Rubiaceae</subject>
        </subj-group>
        <subj-group subj-group-type="scientific_subject">
          <subject>Taxonomy</subject>
        </subj-group>
        <subj-group subj-group-type="geographical_area">
          <subject>Asia</subject>
        </subj-group>
      </article-categories>
      <title-group>
        <article-title>Reappraisal of <italic>Nertera</italic> (Rubiaceae) in Taiwan</article-title>
      </title-group>
      <contrib-group content-type="authors">
        <contrib contrib-type="author" corresp="no">
          <name name-style="western">
            <surname>Chen</surname>
            <given-names>Wei-Chih</given-names>
          </name>
          <xref ref-type="aff" rid="A1">1</xref>
        </contrib>
        <contrib contrib-type="author" corresp="no">
          <name name-style="western">
            <surname>Wang</surname>
            <given-names>Chih-Chiang</given-names>
          </name>
          <xref ref-type="aff" rid="A2">2</xref>
        </contrib>
        <contrib contrib-type="author" corresp="yes">
          <name name-style="western">
            <surname>Chang</surname>
            <given-names>Kun-Cheng</given-names>
          </name>
          <email xlink:type="simple">kcchang@mail.ncyu.edu.tw</email>
          <uri content-type="orcid">https://orcid.org/0000-0001-5807-8546</uri>
          <xref ref-type="aff" rid="A3">3</xref>
        </contrib>
      </contrib-group>
      <aff id="A1">
        <label>1</label>
        <addr-line content-type="verbatim">Graduate Institute of Bioresources, National Pingtung University of Science and Technology., Pingtung, Taiwan</addr-line>
        <institution>Graduate Institute of Bioresources, National Pingtung University of Science and Technology.</institution>
        <addr-line content-type="city">Pingtung</addr-line>
        <country>Taiwan</country>
      </aff>
      <aff id="A2">
        <label>2</label>
        <addr-line content-type="verbatim">Department of Forestry, National Pingtung University of Science and Technology., Pingtung, Taiwan</addr-line>
        <institution>Department of Forestry, National Pingtung University of Science and Technology.</institution>
        <addr-line content-type="city">Pingtung</addr-line>
        <country>Taiwan</country>
      </aff>
      <aff id="A3">
        <label>3</label>
        <addr-line content-type="verbatim">No.300 Syuefu Rd., Chiayi City 60004, Taiwan (R.O.C.), Chiayi, Taiwan</addr-line>
        <institution>No.300 Syuefu Rd., Chiayi City 60004, Taiwan (R.O.C.)</institution>
        <addr-line content-type="city">Chiayi</addr-line>
        <country>Taiwan</country>
      </aff>
      <author-notes>
        <fn fn-type="corresp">
          <p>Corresponding author: Kun-Cheng Chang (<email xlink:type="simple">kcchang@mail.ncyu.edu.tw</email>).</p>
        </fn>
        <fn fn-type="edited-by">
          <p>Academic editor: </p>
        </fn>
      </author-notes>
      <pub-date pub-type="collection">
        <year>2021</year>
      </pub-date>
      <pub-date pub-type="epub">
        <day>27</day>
        <month>05</month>
        <year>2021</year>
      </pub-date>
      <volume>1</volume>
      <uri content-type="arpha" xlink:href="http://openbiodiv.net/2484BDF1-36EE-5057-89C4-1B2A85A4F739">2484BDF1-36EE-5057-89C4-1B2A85A4F739</uri>
      <history>
        <date date-type="received">
          <day>27</day>
          <month>05</month>
          <year>2021</year>
        </date>
        <date date-type="accepted">
          <day>27</day>
          <month>05</month>
          <year>2021</year>
        </date>
      </history>
      <permissions>
        <copyright-statement>Wei-Chih Chen, Chih-Chiang Wang, Kun-Cheng Chang</copyright-statement>
        <license license-type="creative-commons-attribution" xlink:href="http://creativecommons.org/licenses/by/4.0/" xlink:type="simple">
          <license-p>This is an open access preprint distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.</license-p>
        </license>
      </permissions>
      <abstract>
        <label>Abstract</label>
        <p>Only one species in the genus <italic>Nertera</italic>,<italic> N. granadensis</italic>, had been identified in Taiwan. This study discovered that there are two additional species in the genus endemic to Taiwan: <italic>N. nigricarpa</italic> and <italic>N. </italic><italic>taiwanian</italic>. These species are both distinguishable from <italic>N. granadensis</italic> through an examination of their holotype and living individuals. <italic>N. nigricarpa</italic> is characterized by the unapparent or slightly convex leaf venation on the upper surface, spreading leaf margins, purple petals, black fruits, and dark-purple seeds with raised striate. <italic>N. </italic><italic>taiwanian</italic> has leaves with distinctly convex veins on the upper surface, undulated margins, yellowish-green petals, red fruits, and yellow-white seeds without striate.</p>
      </abstract>
    </article-meta>
  </front>
</article>
