Carmen Benítez-Benítez, Pedro Jiménez-Mejías, Santiago Martín-Bravo contributed to the study conception and design of the manuscript. Data preparation was carried out by Carmen Benítez-Benítez, Pedro Jiménez-Mejías, M. Luceño, and Santiago Martín-Bravo. Carmen Benítez-Benítez, Pedro Jiménez-Mejías, and Santiago Martín-Bravo collected plant material. Analyses were carried out by Carmen Benítez-Benítez. The first draft of the manuscript was written by all authors who also commented and contributed on the following versions of the manuscript. All authors contributed to the writing of the final version and approved it.